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Denis Llewellyn Fox (1901-1983), was a chemist who came to Scripps Institution of Oceanography in 1931. During his early years at Scripps, Fox continued his studies of carbon dioxide narcosis and investigated pigments in fishes. His lifelong research in fouling began in 1932 in collaboration with Scripps Director Thomas Wayland Vaughan and chemist Erik Gustaf Moberg. He was associated with the San Diego Zoological Society as their scientific advisor and is remembered for his research on the problem of nutrition in flamingo's to keep their pink color. He is perhaps best known for his two books Animal Biochromes (1953) and Biochromy: Natural Coloration of Living Things (1979). 1936.

Denis Llewellyn Fox (1901-1983), was a chemist who came to Scripps Institution of Oceanography in 1931. During his early y...