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Denis Llewellyn Fox (right) with an unidentified man. Fox's primary research interests were in the pigments of animals, marine, terrestrial and aerial. His studies of these animal biochromes led to important discoveries of both fundamental and practical consequences. For instance, he discovered how to restore color to the fading flamingos of the San Diego Zoo. He had enjoyed the longest tenure of any faculty member of the Scripps Institution of Oceanography--52 years in the discipline of marine biochemistry. Circa 1947.